Field Guide to Intermountain Cyperaceae (excluding Carex) › Cladium

Cladium californicum (S. Watson) O’Neill

California sawgrass, Sawgrass

Figure 9

Synonyms:

Cladium mariscus (L.) Pohl var. californicum S. Watson, Mariscus californicus (S. Watson) Fernald

Plant habit:

Coarse perennial forming dense clumps; rhizomes stout, 20 cm long, 10 mm in diameter; culms subterete to subtriangular, 1–2 m tall, 5–10 mm thick.

Leaves:

Cauline; ligules absent, blades flat to broadly V-shaped, serrated with cartilaginous teeth along the margins and abaxial side of the midvein, 1–2 m long, 7–10 mm wide.

Involucral bracts:

Three or more, leaf-like, spreading.

Inflorescences:

Terminal and lateral, panicle-like, purplish brown, 30–50 cm long with flexuous branches (to 3rd and sometimes 4th order); peduncles at least partially covered by the leaf sheaths; spikelets numerous, mostly in groups of (3) 4–6, ellipsoid to lanceolate with acute apices, reddish brown, to 3 mm long.

Scales:

Few, spirally arranged, ovate to oblong-lanceolate with a distinct midvein, reddish-brown, 2.5–3 mm long, 2 mm wide, lowest scale empty, middle scale empty or subtending staminate flowers, terminal scale subtending a perfect flower.

Flowers:

Perfect and staminate; perianth lacking; stamens 2, anthers 2 mm long, connective apices 0.1 mm; styles trifid, 1–1.5 mm long; stigmas 1–1.5 mm long.

Achenes:

Ovoid, barely trigonous, the base truncate and the apex acute, smooth or irregularly rugulose, glossy purplish brown, 1.5–2 mm long, 1 mm wide; the persistent style base forms a beak 0.2 mm long.

Fruiting: spring—summer.
Habitat and distribution:

Alkaline marshes, swamps, and springs from sea level to 2000 m elevation. Rare in the Intermountain Region. Occurs in Arizona, California, southern Nevada, southern Utah, Texas, and Mexico. Most Utah populations were inundated by Lake Powell.

Cladium californicum: photographs of a densely branched inflorescence and a mass of achenes. Drawings show the flowering stem, a spikelet cluster and enlarged spikelet and achene.
Figure 9. Cladium californicum. (A) Inflorescence, (B) achenes. Photos: (A) BLM SOS_2003_NV-052. (B) J. McDonald (California Botanic Garden). Drawings: a-c: E.E. Reid (Tucker 2012), d: Y. Wilson-Ramsey (Tucker 2002a).

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