Field Guide to Intermountain Cyperaceae (excluding Carex) › Dulichium

Dulichium arundinaceum (L.) Britton
var. arundinaceum

Three-way sedge

Figure 21

Synonyms:

Cyperus arundinaceus L.

Plant habit:

Perennial; rhizomes long, 3–5 mm thick; culms 2–14, arising singly, stout, erect, cylindric, stout, jointed with hollow internodes, leafy, 2–10 dm tall, 4 mm thick.

Leaves:

Numerous, well distributed along the stem; lower 2–14 leaves firm, bladeless, green sheaths with brown, hyaline margins at the mouth, sheaths not splitting; upper 3-15 leaves spreading or ascending with flat, leathery, glabrous 3-ranked grasslike blades with acute tips, 4–15 cm long, (1.5) 3.5–8 mm wide; distal 5–20 leaves are bracts.

Inflorescences:

6–30 cm long, consisting of 1-17 spikes, about 2-5 cm long in sheath axils; spikelets 5-20 per spike, 2-ranked, flat, ascending to spreading, linear-lanceolate, greenish to brownish, 15–30 mm long, 1.2–2.5 mm wide; rachilla hyaline winged, readily disarticulating with the scale at maturity.

Scales:

4–8 per spike, 2-ranked, decurrent, lanceolate, acute to acuminate, prominently many-nerved, light green becoming pale, hyaline along the margins, spongy at the base, (3) 5–9.5 mm long, about 2 mm wide.

Flowers:

Perfect, sessile, borne singly in the scale axils; perianth bristles 6–9, rigid, retrorsely barbellate, about 4–8 mm long, from shorter than the achene to much longer; stamens 3, longer than the perianth bristles; anthers 3-4 mm; stigmas 2, deeply branched, much exerted from the scale.

Achenes:

Narrowly ellipsoid, flattened or somewhat planoconvex with a short, stout stipe, yellowish brown, 2.5–4 mm long (excluding the 1 mm style base), about 1 mm wide; the persistent, undivided basal portion of the style forms a linear beak.

Fruiting: July–October.
Habitat and distribution:

Marshy shores, wet meadows, bogs, pond margins, and ditch banks from sea level to 700 m. Distributed from southwestern British Columbia to California and in eastern North America. Rare in the Intermountain Region, known from Valley County, Idaho and east central California.

Dulichium arundinaceum var. arundinaceum: photographs of a narrow, branched inflorescence, scales and achenes retaining long styles and stigmas; two fruits also retain bristles. Drawings show the leafy culm, leaf sheaths and spikelet arrangement.
Figure 21. Dulichium arundinaceum var. arundinaceum. (A) Inflorescence, (B) scales, (C) achenes with styles and stigmas attached, center and right with perianth bristles attached. Photos: E.G. Hurd. Drawings: J.R. Janish (Hitchcock et al. 1969).

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