Field Guide to Intermountain Cyperaceae (excluding Carex) › Cyperus

Cyperus squarrosus L.

Bearded flatsedge, Awned flatsedge, Bearded nutgrass

Figure 18

Synonyms:

Cyperus aristatus Rottb., C. inflexus Muhlenberg

Plant habit:

Very small tufted annual with fibrous roots; fresh herbage sweet, celery-scented; dry herbage clover scented; culms 1–20, slender, sharply triangular, smooth, unequally ribbed or striated, 0.3–1.5 dm tall, 0.5–2 mm wide.

Leaves:

Few, all borne near the base, flat to V-shaped, keeled distally, light green, shorter to longer than the culms, 2–10 cm long, 0.5–2.5 mm wide; sheaths of older leaves reddish brown.

Involucral bracts:

Leaflike, 1–6, elongate, unequal, 1–15 cm long, some or all surpassing the inflorescence.

Inflorescences:

1–4 capitate clusters of spikelets, 5–15 mm wide; terminal cluster sessile, others (if present) on slender rays up to 3 cm long; spikelets 1–30 in a cluster, lanceolate to oblong, somewhat compressed, greenish, yellowish, or reddish brown, 4–10 mm long, 1–2.2 mm wide.

Rachilla:

Straight, wingless, persistent.

Scales:

4–20 per spikelet, somewhat closely set, oblong-lanceolate, 5–11 nerved (prominent when dry), slenderly bristle tipped, prominently spreading or recurved (prominent when dry), greenish or yellowish to reddish brown, midrib commonly greenish or paler, 1–2 mm long including the awn, 0.4–0.8 mm wide, deciduous.

Flowers:

Perfect; perianth lacking; stamens 1; style trifid.

Achenes:

Narrowly obovoid or oblong, trigonous, obtusely angled, papillose, stipitate, apex truncate and apiculate, light brown or reddish brown to blackish, 0.6–1.0 mm long, 0.3–0.5 mm wide; stipe 0.06–1.0 mm.

Fruiting: June–November.
Habitat and distribution:

Seeps, springs, disturbed areas in wet meadows and wetlands, flood plains, drying margins of ponds and lakes, and river sandbars from sea level to 2500 m elevation. Occurs throughout much of the continental United States and Mexico, including scattered locations across the Intermountain Region.

Cyperus squarrosus: photographs of culms with rounded spikelet clusters, a spikelet with outward-curving scale tips, detached scales and achenes. Drawings show the tufted plant, spikelet and fruit details.
Figure 18. Cyperus squarrosus. (A) Plant with inflorescences, (B) spikelet, (C) scales, (D) achenes. Photos: E.G. Hurd. Drawings: J.R. Janish (Hitchcock et al. 1969).

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