Field Guide to Intermountain Cyperaceae (excluding Carex) › Schoenoplectus

Schoenoplectus californicus (C.A. Meyer) Soják

California bulrush, Giant bulrush, Tule

Figure 55

Synonyms:

Scirpus californicus (C.A. Meyer) Steudel

Plant habit:

Stout perennial, forming large colonies from long, robust rhizomes 10–15 mm in diameter; culms bluntly triangular throughout, or nearly cylindric and becoming obscurely triangular near the inflorescence, firm and not easily crushed between the fingers, 1–4 m tall, 4–10 mm thick.

Leaves:

3–4, basal; sheaths one-sixth of culm length, fronts ventrally hyaline and splitting leaving coarse fibers; blades 0–1, nearly flat, smooth, up to 2 cm long and 2 mm wide, about half the sheath length.

Involucral bracts:

Main bract usually erect, stiff, margins often scabrous, 1–8 cm long, sometimes resembling a continuation of the culm, shorter than the inflorescence.

Inflorescences:

Panicle-like and open with third level branching; branches arched and flexible, 21 cm long; spikelets 25–150 or more, solitary or in clusters of 2 or more at the branch tips, 5–11 mm long, about 3 mm thick.

Scales:

Oval, orangish brown with numerous short, linear reddish spots, 2.5 mm long, 2 mm wide, smooth except for the thick, sparsely scabrous midrib, margins minutely ciliate, apex rounded, shallowly notched 0.1–0.2 mm deep; awn not contorted, bent outward, about 0.3–0.5 mm long.

Flowers:

Perfect: perianth bristles 2–4, strap-shaped or trigonous with blunt tips, edges fringed, dark reddish brown, equaling the achene; stamens 3; styles all bifid.

Achenes:

Ovoid to obovoid, lenticular, smooth, gray-brown, 1.8–2.2 mm long (including the 0.2 mm long beak), 1.3 mm wide.

Fruiting: spring—summer.
Habitat and distribution:

Fresh to brackish water, marshes and shorelines from sea level to about 1400 m elevation. Often emergent and salinity tolerant. In the Intermountain Region, known only from along the Colorado River in southernmost Nevada. California to Florida; Mexico to South America, West Indies, Pacific Islands.

Similar species:

Easily confused with Schoenoplectus tabernaemontani and S. acutus, both of which have completely cylindric culms and (4) 6–8 perianth bristles with sharp-pointed hairs.

Schoenoplectus californicus: photographs of a branched inflorescence and three enlarged achenes, the central one retaining bristles. Drawings show the upper culm and inflorescence, a scale, a bristled achene and its cross section.
Figure 55. Schoenoplectus californicus. (A) Inflorescence, (B) achenes. Photos: (A) G. Carr (Oregon Flora), (B) C. Ritchie (USDA ARS Plants.usda.gov). Drawings: S.A. Reznicek (Smith 2002c).

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